Timothy W. Sipe

Department of Biology
Franklin & Marshall College

Lancaster, PA 17604
717-291-4188 (office)
717-291-3966 (lab)
717-358-4548 (fax)
Timothy.Sipe@fandm.edu

Further Information: http://www.fandm.edu/Departments/Biology/People/Sipe/Sipe.html

EDUCATION

B. A. 1978 Wabash College, Crawfordsville, IN (Biology)
M.S. 1980. University of Tennessee. Knoxville, TN (Ecology)
PhD. Candidate 1982-1984 University of Illinois. Urbana, IL (Graduate Program in Ecology)
PhD. 1990 Harvard University. Cambridge, MA (Ecology)

POSITIONS

1997-present Associate Professor of Biology, Franklin & Marshall College, Lancaster, PA
1990-97 Assistant/Associate Professor of Biology, Gustavus Adolphus College, St. Peter, MN
1984-90 Graduate Research Assistant, Harvard University, Cambridge, MA
1982-84 Research Assistant, Illinois Natural History Survey, Champaign, IL
1982 Visiting Instructor, Wabash College, Crawfordsville, IN
1980-81 Ecologist, National Coastal Ecosystems Team, U.S. Fish & Wildlife Service, Slidell, LA

RESEARCH INTERESTS

Plant physiological ecology; Plant community ecology; Forest microenvironmental patterns, especially spatial-temporal heterogeneity of multiple factors; Ecosystem disturbance and recovery

RECENT RESEARCH PROJECTS & SITES

• Measurement and Modeling of Photosynthetic Responses by Temperate Forest Herbs and Tree Seedlings to Nearground Enriched Carbon Dioxide and Sunfleck Regimes
  
  • 1998-present, Harvard Forest Long-Term Ecological Research (LTER) Site, central MA
  • Funded by NSF CRUI and RUI Programs
  • Quantification of nearground enriched CO2 (NEC) profiles and their environmental controllers, nearground sunfleck regimes, and the combined effects of NEC and soil moisture on photosynthesis during sunflecks by seedlings of three maple species (Acer saccharum, A. rubrum, A. pensylvanicum) and by three species of herbs with monolayered canopies at different heights (Clintonia borealis, Medeola virginiana, Aralia nudicaulis).
• The Impact of Land-Use Legacies on Forest Soil-Plant-Atmosphere Interactions in Central New England
  • 1995-present, Harvard Forest Long-Term Ecological Research (LTER) Site, central MA
  • Funded by NSF CRUI Program.
  • Three colleges (Franklin & Marshall, Allegheny, Mt. Union), one faculty member and nine students from each institution, plus three research technicians. Sixteen hypotheses on long term effects of 18th and 19th century agricultural land uses (plowed, pastured, permanent woodlot) on forest structure and processes, in five main categories: (1) stocks and fluxes of nitrogen and carbon; (2) forest productivity and litter chemistry, (3) spatial-temporal variation in microclimatic and soil resource factors; (4) ecophysiology of tree seedlings and herbs; (5) community composition, spatial patterns, and forest regeneration
• Forest Composition, Soil Properties, and Tree Seedling Performance in Response to Bedrock Variation in Lancaster County, Pennsylvania
  • 1998-present.
  • Funded by Beckman Scholarship Program, Biology Department Independent Study Funds, and Competitive Research Grants to Individual Students.
  • Fifteen study sites with permanent plots in mature, mixed-hardwood forests on public and private land, with three sites each on limestone, schist, shale, sandstone, and quartzite.

PUBLICATIONS (last 10 years, * = Undergraduate Co-Author)

Catovsky, S., R. Crabtree, T. Sipe, G. Carlton, S. Bassow, and F. Bazzaz. 2003. Experimental approaches to understanding forest regeneration. In Foster, D.R. and J. Aber (eds.), Forests in Time: The Environmental Consequences of 1000 Years of Change in New England. Yale University Press, New Haven, Connecticut.

Sipe, T.W. and F.A. Bazzaz. 2001. Shoot damage effects on regeneration of maples (Acer) across an understory-gap gradient. Journal of Ecology 89(5):761-773.

DeLucia, E.H., T.W. Sipe, J.A. Herrick*, and H. Maherali. 1998. Sapling biomass allocation and growth in the understory of a deciduous hardwood forest. American Journal of Botany 85(7):955 963.

Maherali, H., E.H. DeLucia, and T.W. Sipe. 1997. Hydraulic adjustment of maple saplings to canopy gap formation. Oecologia 112:472-480.

Sipe, T.W. and A.R. Linnerooth*. 1995. Intraspecific variation in samara morphology and flight behavior in Acer saccharinum. American Journal of Botany 82:1412-1419.

Sipe, T.W. and F.A. Bazzaz. 1995. Gap partitioning among maples (Acer) in central New England: survival and growth. Ecology 76:1587-1602.

Sipe, T.W. and F.A. Bazzaz. 1994. Gap partitioning among maples (Acer) in central New England: shoot architecture and photosynthesis. Ecology 75:2318-2332.

RECENT PRESENTATIONS (last 5 years, * = Undergraduate Co-Author)

Burton, P.J., C.M. Burton, T.W. Sipe, and D.E. Hibbs. 2003. Tree regeneration and understory vegetation in an eastern hardwood forest twenty years after creation of small forest openings. Annual Ecology Symposium, Harvard Forest, Harvard University, Petersham, MA, February 2003.

Sipe, T.W., C.S. Pike, and J.R. Richardson. 2001. Integrating the study of photosynthesis across multiple scales in the biology majors curriculum. Ecological Society of America, August, 2001, Madison, WI.

Sipe, T.W., C.A. McClaugherty, and R. D. Bowden. 2001. The effects of nearground enriched carbon dioxide on photosynthesis in forests with different land use legacies. Ecological Society of America, August, 2001, Madison, WI.

McClaugherty, C.A., T. W. Sipe and R. D. Bowden. 2000. Near ground light and carbon resource availabilities in temperate forests with contrasting historical land uses. American Association for the Advancement of Science, February 2000, Washington, DC. Published in abstracts of meeting A75.

Kunkle*, J.M, T.W. Sipe, R.D. Bowden and C.M. McClaugherty. 1999. Growth rates and biomass allocation of tree seedlings across forests with three different land-use legacies in central Massachusetts. Ecological Society of America, August, 1999, Spokane, WA.

Bixler*, K.S., T.W. Sipe, C.M. McClaugherty and R.D. Bowden. 1999. Spatial variation of sunfleck regimes in forests with different land-use legacies in central Massachusetts. Ecological Society of America, August, 1999, Spokane, WA.

Bowden, R.D., C. McClaugherty, T. Sipe, K. Clark*, C. Giffen*, C. Meredith*, S. Stewart.* 1998. Legacies of agricultural activities in New England forest soils. Proceedings of the Ninth North American Forest Soils Conference, August 9-14, 1988, Tahoe City, CA